Taxonomy and phylogeny of the species of the weevil Miarus Schoenherr, 1826 (Coleoptera: Curculionidae, Curculioninae)

The Palaearctic genus Miarus SCHONHERR, 1826 (Coleoptera: Curculionidae, Curculioninae) is revised. Nineteen species are recognized as valid. Miarus ursinus ABEILLE ssp. maroccanus SOLARI, 1947 is considered as distinct species. The following new synonymies are proposed: M. abnormis SOLARI, 1947 (= M. zoufali SOLARI, 1947 syn.n.), M. ajugae (HERBST, 1795) (= M. portae SOLARI ssp. confusus ROUDIER, 1966 syn.n., = M. thuleus KANGAS, 1980 syn.n.), M. campanulae (LINNAEUS, 1767) (= M. moroi SOLARI, 1947 syn.n.), M. dentiventris REITTER, 1907 (= M. armenus IABLOKOFF-KHNZORIAN, 1967 syn.n.), M. maroccanus SOLARI, 1947 (= M. italicus FRANZ ssp. maroccanus FRANZ, 1947 syn.n.), M. monticola PETRI, 1912 (= M. fennicus KANGAS, 1978 syn.n.), M. simplex SOLARI, 1947 (= M. alzonae SOLARI, 1947 syn.n., = M. portae SOLARI, 1947 syn.n.). The lectotypes of the following taxa are designated: Curculio ajugae HERBST, 1795, Miarus abeillei DESBROCHERS DES LOGES, 1893, M. abnormis SOLARI, 1947, M. alzonae SOLARI, 1947, M. araxis REITTER, 1907, M. binaghii SOLARI, 1947, M. brevirostris SOLARI, 1947, M. dentiventris REITTER, 1907, M frigidus FRANZ, 1947, M. horni FRANZ, 1947, M. italicus FRANZ, 1947, M. longicollis SOLARI, 1947, M. moroi SOLARI, 1947, M. muelleri SOLARI, 1947, M phyteumatis FRANZ, 1947, M. phyteumatis imitator FRANZ, 1947, M. portae SOLARI, 1947, M. rotundicollis DESBROCHERS DES LOGES, 1893, M. simplex SOLARI, 1947, M. stoeckleini FRANZ, 1947, M. subseriatus SOLARI, 1947, M. ursinus maroccanus SOLARI, 1947, M. zoufali SOLARI, 1947. A key to the species and descriptions of diagnostic characters, biological and distributional notes, as well as illustrations of male genitalia of all species are provided. Moreover a cladistic analysis was performed to reconstruct the phylogenetic relationships of the species of this genus. Evolutionary trends of some characters are discussed.