Coniocleonus

genus Coniocleonus Motschulsky, 1860: c. 540 type species Cleonus carinirostris Gyllenhal, 1834 (= Cleonus schoenherri Gebler, 1829)

subgenus Angarocleonus Arzanov, 2006c: 115 type species Cleonus cineritius Gyllenhal, 1834
astragali Ter-Minasyan & Korotyaev, 1977: 825 !  TL: Altai  ASIA: RU*
cinerascens Hochhuth, 1851: 32 (CleonusTL: “Süd-Ost Sibirien”  ASIA: MN2,# RU*
cineritius Gyllenhal, 1834: 177 (Cleonus) !  TL: Sibiria orientali”  ASIA: IR2,35 KP21 MN2,# RU*
elisabethae Ter-Minasyan & Korotyaev, 1977: 831  TL: Burjatia, Amur, Mongolia  ASIA: MN* RU*
ferrugineus Fåhraeus, 1842: 24 (Cleonus) !  TL: “Tataria magna”  ASIA: CN(GS2) KZ* MN2,59,# RU2,15,#
      hexastichus Boheman, 1842: 25 (Cleonus) !  TL: “Irkutsk”  ASIA: RU*
parshus Anderson, 1987: 461 (Stephanocleonus) !  TL: “Chalk river, Ontario”  NAM: CA* US(CT43 IA43 MA43 ME43 MI43 MN43 MT43 NE43 NM43 SD43 TN43 VT43
plumbeus LeConte, 1876a: 146 (Stephanocleonus)  TL: “Lake Superior”  NAM: CA*
zherichini Ter-Minasyan & Korotyaev, 1977: 829 !  TL: “Tajmirsk ochr.”ASIA: RU*  NAM: CA41
stenothorax Anderson, 1987: 465 (Stephanocleonus)  TL: “Bluefish Caves, 67°08’N 140°48’W”  NAM: CA*

subgenus Augustecleonus Arzanov, 2006b: 115 type species Curculio nebulosus Linnaeus, 1758
hollbergii Fåhraeus, 1842: 10 (Cleonus) !  TL: “Saisan lake”  ASIA: KZ* RU2,15  EUR: AT1,# BY13 CZ19,32,# DE1,# FR1# HU# IT1,# LT12 LV9 PL1,7 RU1,# SE1,54 UA23  EUR(uncertain presence, see remarks) BA1 BE1 BG1,52 CH1 DK1 ES1 FI1 NL1 NO1 RO1 SK1
nebulosus Linnaeus, 1758: 385 (Curculio) !  TL: “Europa”  EUR: BE1,11 BG1,52 BY13 CZ1,19,# DE1,# DK1,§ ES1,48 FI1 FR1 GB1,16,# IT1,37 LT12 LU39 LV9,# NL1 PL1,7 RU1 SE1,54,§ SI57 SK19  Photo of the type: see "The Linnean Collections, LINN 7737".
      carinatus DeGeer, 1775: 241 (Curculio) [Homonym]  TL: no distribution reported
      ericeti Dufour, 1843: 80 (Cleonis)  TL: “Vallée d’Ossau”  EUR: FR*
      guttulatus Gyllenhal, 1834: 178 (Cleonus) !  TL: no distribution reported
      iners Scopoli, 1763: 24 (Curculio) TL: “Carniolia”
      incanescens Panzer, 1795: 322 (Curculio)  TL: “Germania”  EUR: DE*
      lethierryi Chevrolat, 1873: 24 (Plagiographus) !  TL: “Renosa” (Reynosa, Spain)  EUR: ES*
turbatus Fåhraeus, 1842: 9 (Cleonus) !  TL: “Europa borealis et meridionalis”  EUR: AT1 BE1,14 BG1 CH38 CZ19,32,# DE1,# DK1 ES48 FI1 FR1,# GB16,# HU1,# IT37 LU39 LV9,# PL1,7,# RU1,2,# SE1,# SI# SK19,# UA1,#
      glaucus Fabricius, 1787: 114 (Curculio) [Homonym]  TL: “Kilia”  EUR: UA*
      glaucus nubilus Desbrochers des Loges, 1905: 34 (Cleonus) [Homonym]  TL: “Savoie”  EUR: FR*
      hubenthali Krausse, 1928: 24 (Cleonus)  TL: Germany  EUR: DE*
      nebulosus quadriguttulus Desbrochers des Loges, 1891b: 29 (Bothynoderes)  TL: Central France  EUR: FR*
      obesulus Desbrochers des Loges, 1893: 6 (Cleonus)  TL: “Espagne”  EUR: ES*
      sulcatus Goeze, 1777: 381 (Curculio) [Homonym]  TL: no distribution reported
      sulcatus Geoffroy, 1785: 117 (Curculio) [Homonym]  TL: “Paris”  EUR: FR*
      sulcatus Gmelin, 1790: 1758 (Curculio) [Homonym]  TL: “Gallia”  EUR: FR*

subgenus Borisocleonus Arzanov, 2006: 64 type species Lixus mesopotamicus Olivier, 1807
mesopotamicus Olivier, 1807: 264 (Lixus) !  TL: “Alah, Mesopotamia”  ASIA: AF27,58,§ AM27,# AZ# CY8 IN45,# IQ* IR27,35 KZ# PK50 SY35
      lederi Faust, 1891a: 398 (Neocleonus)  TL: “Armenischen Gebirge; Araxes-Tahl”  ASIA: AM*
vittiger Fåhraeus, 1842: 11 (CleonusTL: “Tauria”  AFR: EG5  ASIA: CY1,8,# IL17 IQ35 IR35 RU2,15 SY35,# TR26,#  EUR: GR1,28,# RU1,2 UA*
      frenatus Reiche & Saulcy, 1854: n. 545 (Cleonus) (nomen nudum)  TL: “Syria”  ASIA: SY*

subgenus Coniocleonus Motschulsky, 1860: c. 540 type species Cleonus carinirostris Gyllenhal, 1834 (= Cleonus schoenherri Gebler, 1829)
alpinus Gebler, 1833: 294 (Cleonus) !  TL: “Altai”  ASIA: KZ*
schoenherri Gebler, 1829: 162 (Cleonus) !  TL: “Barnaul”  ASIA: KP21 MN# RU*
      carinirostris Gyllenhal, 1834: 176 (Cleonus) !  TL: “Sibiria orientalis”  ASIA: RU*
vinokurovi Ter-Minasyan & Korotyaev, 1977: 826 !  TL: Amur, Yakutia  ASIA: RU*

subgenus Plagiographus Chevrolat, 1869: 76 type species Stephanocleonus saintpierrei Chevrolat, 1869 (= Cleonus variolosus Wollaston, 1864)
amori Marseul, 1868: 199 (StephanocleonusTL: no indication reported  EUR: ES1,48,# PT1,#
      gaditanus Chevrolat, 1869: 76 (Stephanocleonus)  TL: “Cadix”  EUR: ES*
cicatricosus Hoppe, 1795: 61 (CurculioTL: “Erlangen”  AFR: MA4,# TN3  ASIA: "Caucasus"2 "Transcaucasia"2 AM# CY1,8 GE# IQ# IR35,49 TM# TR#  EUR: BA1 CH38 CZ32 DE* ES1,48,# FR1 GR1,28,# HR1 HU1 IT1,37 LU39 MK# PL1,7 RO1 RS1,31 SI1 SK1,19,# TR#
      arciferus Chevrolat, 1874: 158 (Plagiographus) !  TL: “Aranjuez, Grenoble”  EUR: ES*  FR*
      callosus Bach, 1854: 230 (Cleonus)  TL: “Boppard”  EUR: DE*
      ericae Fåhraeus, 1842: 36 (Cleonus) !  TL: no indication reported
      spinosus Ponza, 1805: 63 (Curculio) [Homonym]  TL: Valle Varaita  EUR: IT*
crinipes Fåhraeus, 1842: 35 (Cleonus) !  TL: “Caucasus”  AFR: DZ35,# EG#,35 LY10,# MA4,# TN35,#ASIA: "Transcaucasia"2 AM# AZ# CY1,8,# GE# IL17,# IR35,# JO18 KZ29 LB# SY6 TR#  EUR: ES1,48,# RU1,#
      crinipes signifer Chevrolat, 1876: cxlvii (Plagiographus) !  TL: “Syrie”  ASIA: SY*
      crinipes uniformis De La Fuente, 1912: 363  TL: “Cabeza-parda, Pozuelo”  EUR: ES*
excoriatus Gyllenhal, 1834: 194 (Cleonus) !  TL: “Gallia boreali, Lusitania”  AFR: DZ# EG5,# ES(Canary Islands)1,46,# LY10,# MA4,# TN3,#  ASIA: "Caucasus"2 CY1,8,# EG(Sinai)5 IL17,# IQ36,# IR35,51 JO18,# PK# PS# SY6,# TR42 YE#  EUR: AT1,# BE1,14 BG1,52 CH38 CZ32 DE1 ES1,48,# FR* GI53,# GR1,28,# HU1 IT1,37,# MK# MT1,22,# PT* RU1,2 SE1 SK1 UA1,23,#
      albirostris Chevrolat, 1873: 23 (Plagiographus) !  AFR: EG*
      lacunosus Gyllenhal, 1834: 194 (Cleonus) !  AFR: MA*
      sulcicollis Fåhraeus, 1842: 34 (Cleonus)  EUR: IT(Sicily)*
fastigiatus Erichson, 1841: 187 (Cleonus)  TL: “Algier”  AFR: DZ* MA4,#
graellsi Chevrolat, 1873: 25 (Plagiographus) !  TL: “Madrid, Reinosa”  EUR: ES* PT#
megalographus Fåhraeus, 1842: 33 (Cleonus) !  TL: “Sicilia”  AFR: MA#  EUR: ES48 FR1 GR# IT* MK# MT#
nigrosuturatus Goeze, 1777: 381 (CurculioTL: distribution not indicated  AFR: DZ# EG5 LY10 MA4,# TN3,#  ASIA: AF§ AM# AZ33,# CY1,8,# GE# IL17,# IN2 IQ# IR2,35,# JO18 KG# KZ# LB# PK# PS# RU2,# SY6# TJ# TM2# TR26,# UZ2,59,#  EUR: AL# AT1,# BE1 BG1,# CH38 CZ1,19 DE1 ES1,# FR1 GR1,28 HR1,34 HU1,20# IT1,37 MD1,40 PT1,# RO24,# RS31,# RU1,2,# TR1,25 UA1,23
      obliquus Fabricius, 1792: 460 (Curculio)  TL: “Germania”  EUR: DE*
      leucomelanus Fåhraeus, 1842: 32 (Cleonus)  TL: “Ind. or.”
      paraplecticus Geoffroy, 1785: 117 (Curculio) [Homonym]  TL: “Paris”  EUR: FR*
planidorsis Fairmaire, 1874: 388 (Cleonus) !  TL: “Souff”  AFR: DZ* EG#,5 LY10,56,# MA# TN56,#  ASIA: CY1,44 IL# JO# SA#  EUR: ES48
pseudobliquus J. Müller, 1921: 75 (CleonusTL: “Dalmatien, Süd-Istriem, Griechenland, Mazedonien”  ASIA: IQ36 TR#  EUR: BA1 BG# GR* HR* IT1,37,# MK* MT22 RO47 RS1,31 RU(Crimea)# SI#F TR# UA#
riffensis Fairmaire, 1884: 446 (Cleonus) !  TL: “Casablanca”  AFR: MA*  EUR: ES48,# GI#
      fasciculosus Reitter, 1890: 157 (Plagiographus)  TL: “Andalusien”  EUR: ES*
sejunctus Faust, 1904: 253 (Stephanocleonus) !  TL: “Huascar”  EUR: ES*
      vasquezi Desbrochers des Loges, 1904: 106 (Cleonus)  TL: Toledo  EUR: ES*
tabidus Latreille, 1804: 122 (CurculioAFR: ES(Canary Islands)1 MA4,#EUR: ES48,# FR1,# GR1,28 IT1,37,# PT1
      montalbicus Costa, 1885: 26 (Cleonus)  TL: Sardegna  EUR: IT*
      pelletii Fairmaire, 1859a: lii (Cleonus)  TL: “Béziers”  EUR: FR*
variolosus Wollaston, 1864: 323 (CleonusTL: “Fuerteventura”  AFR: DZ56,# EG5,# ES(Canary Islands)* LY10,# MA4,56 TN3,#  ASIA: IL17,#  EUR: ES1,48,# IT(Sicilia)1,37
      scabrithorax Desbrochers, 1898a: 14 (Gonocleonus) TL: Algérie  AFR: DZ*
      bonnairei Fairmaire, 1883: xlvi (Cleonus) !  TL: Algérie  AFR: DZ*
      saintpierreorum Chevrolat, 1869: 76 (Plagiographus) !  TL: “Oran”  AFR: DZ*

References: 1: Fauna Europaea 2016; 2: Ter-Minasyan 1988a; 3: Normand 1937; 4: Kocher 1961; 5: Alfieri 1976; 6: Weill & al. 2011; 7: PolBIN; 8: Alziar 2010; 9: Balalaikins & Bukejs 2011; 10: Colonnelli 2013; 11: Delbol 2013; 12: Tamutis & al. 2011; 13: Alexandrovitch & al. 1996; 14: Delbol 2014; 15: Legalov 2010; 16: Duff 2012; 17: Friedmann (pers. comm.); 18: Katbeh-Bader 2008; 19: Benedikt & al. 2010; 20: Podlussany & Zoltán 2008; 21: Hong & Korotyaev 2002; 22: Mifsud & Colonnelli 2010; 23: Mazur 2002; 24: Procheş 1999; 25: Avgın & Colonnelli 2011; 26: Fremuth 1982; 27: Voss 1959; 28: Bahr & al. 2016; 29: Bajtenov 1974; 30: Hua 2002; 31: Pešić & Mesaroš 2012; 32: Stejskal & Trnka 2013; 33: Tozlu & al. 2005; 34: Pešić S. 2003; 35: Legalov & al. 2010; 36: Derwesh 1965; 37: Abbazzi & Maggini 2009; 38: Germann 2010; 39: Braunert 2009; 40: Bacal & al. 2013; 41: Bousquet & al. 2013; 42: Sert & Çağatay 1999; 43: Anderson 1987; 44: Srour 2013; 45: Faust 1904; 46: Machado & Oromí 2000; 47: Podlussâni & Kocs 2013; 48: Alonso-Zarazaga 2002; 49: Ghahari & Colonnelli 2012; 50: Ahmed & Legalov 2015; 51: Modarres Awal 1997; 52: Angelov 1978; 53: Perez & Bensusan 2016; 54: Rassi & al. 2015; 55: Stüben 2016; 56: Bedel 1909; 57: Alonso-Zarazaga & al. 2016; 58: Ter-Minasyan 1972d; 59: Heyden 1895; §: Entomological Museum of Lund University;

Key: Faust 1904; Ter-Minasyan 1988a (Russia); Skuhrovec & al. 2014 (West Palaearctic)

Remarks. Coniocleonus hollbergii (Fåhraeus, 1842) and C. turbatus (Fåhraeus, 1842). Citations for these species in regional faunas and checklists, including Fauna Europea, are uncertain due to misconcepts and misidentifications. Cleonus hollbergii Fåhraeus, 1842 was described from Kazakhstan (Zaisan lake); the application of this name to the European species is uncertain and only countries for which controlled documentation is available are listed. Countries with undocumented citations of C. hollbergii are reported in italics: in most cases they are likely to refer to C. turbatus.

Reiche & Saulcy (1854) listed a Cleonus frenatus from Syria, that was considered a synonym of Coniocleonus vittiger in Csiki (1934). However, this is a nomen nudum, listed but not described.

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